Females Heterozygous for the Recessive Second Chromosome Mutations Pn,px,and Sp

Essay

Females heterozygous for the recessive second chromosome mutations pn,px,and sp are mated to a male homozygous for all three mutations.The offspring are as follows: $$\begin{array} { | l | r | } \hline p x s p c n & 1,461 \\\hline p x s p + & 3,497 \\\hline p x + c n & 1 \\\hline p x + + & 11 \\\hline + s p c n & 9 \\\hline + s p + & 0 \\\hline + + c n & 3,482 \\\hline + + + & \underline { 1,539 } \\\hline & 10,000 \\\hline\end{array}$$
What is the genotype of the females that gave rise to these progeny?