Which Methods Did Seymour Benzer Use to Study 20,000 Mutants

Multiple Choice

Which methods did Seymour Benzer use to study 20,000 mutants of the rII region of the T4 bacteriophage?

A) intragenic recombination analysis and deletion- mapping analysis only
B) genetic complementation analysis only
C) intragenic recombination analysis only
D) genetic complementation analysis, intragenic recombination analysis, and deletion- mapping analysis
E) deletion- mapping analysis only