If Archaeopteryx Had Air Sacs, Then Which of Its Features

Multiple Choice

If Archaeopteryx had air sacs, then which of its features would have had the opposite effect on enabling Archaeopteryx to fly long distances from that provided by air sacs?
1) teeth
2) contour feathers
3) wing claws
4) long tail with many vertebrae

A) 1 and 2
B) 1 and 4
C) 2 and 3
D) 1, 2, and 4
E) 1, 3, and 4