Exam 17: Annelids and Allied Taxa
Of what evolutionary significance is the "hydrostatic skeleton"?
The hydrostatic skeleton is of significant evolutionary importance as it represents a fundamental mechanism by which many organisms have achieved movement, support, and structural integrity. This type of skeleton is characterized by a fluid-filled cavity, the coelom, surrounded by muscles. Unlike rigid skeletons, hydrostatic skeletons rely on the pressure of the fluid inside the body cavity to maintain shape and enable movement.
Here are some key evolutionary significances of the hydrostatic skeleton:
1. **Early Evolution of Movement**: The hydrostatic skeleton allowed the earliest multicellular organisms to move in a coordinated manner. By contracting muscles against the fluid-filled cavity, these organisms could change shape and propel themselves through their environment. This was a significant step in the evolution of more complex locomotion.
2. **Support and Structure**: For soft-bodied organisms, the hydrostatic skeleton provides structural support without the need for hard, mineralized tissues. This allows for a wide range of body shapes and sizes, which can be advantageous in different ecological niches.
3. **Adaptability**: Hydrostatic skeletons are highly adaptable. They can change shape easily, allowing organisms to squeeze through tight spaces or burrow into substrates. This flexibility is advantageous for escaping predators, hunting prey, or simply moving through complex environments.
4. **Energy Efficiency**: Hydrostatic skeletons can be energy-efficient because they do not require the organism to maintain a rigid structure at all times. Muscles can relax when not in use, saving energy.
5. **Evolution of Complex Organisms**: The hydrostatic skeleton has been a stepping stone in the evolution of more complex body plans. For example, the ancestors of cephalopods (like squids and octopuses) and annelids (like earthworms) utilized hydrostatic skeletons. These groups have since evolved a variety of sophisticated adaptations, such as the ability to manipulate objects and complex burrowing behaviors.
6. **Diversification**: Hydrostatic skeletons have allowed for a great diversification of life forms, particularly among invertebrates. Many phyla, including Cnidaria (jellyfish), Annelida (segmented worms), and Mollusca (mollusks), rely on this type of skeletal system.
7. **Evolutionary Precursor**: In some lineages, the hydrostatic skeleton may have been a precursor to the development of more rigid internal or external skeletons. For example, the notochord in chordates, a flexible rod-like structure, is considered a type of hydrostatic skeleton that played a role in the evolution of the vertebral column in vertebrates.
In summary, the hydrostatic skeleton has played a crucial role in the evolution of multicellular life, particularly in soft-bodied organisms. It has facilitated movement, provided structural support, allowed for adaptability in various environments, and served as a precursor to more complex skeletal systems. Its significance is evident in the diversity and success of the myriad organisms that utilize this form of skeletal structure.
When earthworms mate,the _______________ secretes mucus to hold them together.
clitellum
The circulatory system in the earthworm
C
Discuss why segmentation is such an important innovation in the Annelida.
In leeches,a cocoon into which fertilized eggs will be deposited is secreted by the __________.
The most recent molecular evidence suggests that the Annelida and Mollusca are more closely related to each other (in the clade Lophotrochozoa)rather than either phylum is to the Arthropoda (clade Ecdysozoa).Do you think 1)it matters that this distinction is made,and 2)should the use of molecular evidence by itself be the sole means of distinguishing the affinities of these groups? Support your argument.
Paired appendages used for locomotion in polychaetes are called ___________________.
The spaces in the coelom of leeches are called the _______________.
Why do zoologists not have a well-accepted theory for the origin of segmentation?
A biologically plausible reason that burrowing earthworms have solid segmental walls but nonburrowing and parasitic annelids often have partial septa is
In the earthworm,food is drawn into the mouth by the pumping action of the _____________.
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