Exam 8: Principles of Development
In nearly all metazoa,"germ layers" is a term that describes
C
The abnormal entrance of more than one sperm into an egg,normally prevented by the fertilization membrane,is called _______________.
polyspermy
If yolk is primarily food storage for the developing embryo,why is it so central to understanding cleavage?
The yolk within an egg cell is indeed a source of nutrients for the developing embryo, but it also plays a crucial role in the process of cleavage, which is the series of rapid cell divisions that occur after fertilization. Understanding the relationship between yolk and cleavage is important for several reasons:
1. **Yolk Distribution and Cleavage Patterns**: The amount and distribution of yolk in the egg can affect the pattern of cleavage. In eggs with little yolk (isolecithal eggs), such as those of mammals, cleavage is typically holoblastic, meaning that the entire egg divides completely with each cleavage event. In contrast, in eggs with a large amount of yolk (telolecithal eggs), such as those of birds and reptiles, cleavage is meroblastic, meaning that cell division occurs only in the yolk-free cytoplasmic region of the egg. This is because the dense yolk inhibits the cleavage furrow from passing through it.
2. **Rate of Cell Division**: The presence of yolk can slow down the rate of cell division in the areas where it is concentrated. Cells that contain less yolk divide more rapidly than those with more yolk. This leads to an uneven rate of division and can result in different cell sizes, which is important for the future differentiation of cells into various tissues and organs.
3. **Embryonic Axes Formation**: The distribution of yolk within the egg can influence the establishment of the embryonic axes (anterior-posterior, dorsal-ventral, left-right). As cells divide and interact with the yolk and each other, they begin to establish the body plan of the developing organism.
4. **Nutrient Provision**: The yolk not only serves as a nutrient reserve but also influences the timing of gene expression and metabolic activity in the developing embryo. As cells cleave and form the blastula, they will eventually need to access the yolk's resources for further growth and development.
5. **Cell Fate Determination**: The position of cells relative to the yolk can affect their developmental fate. In some species, the yolk contains determinants that influence the developmental pathway of the cells that inherit them during cleavage.
6. **Evolutionary Adaptations**: The adaptations in cleavage patterns in response to yolk distribution reflect the evolutionary pressures on different species. For example, the large yolk of bird eggs supports the development of the embryo on land, without the need for additional water or nutrients from the environment.
In summary, the yolk's role in providing nutrition is closely tied to the mechanics and regulation of cleavage. Its influence on the rate, pattern, and symmetry of cell division, as well as its impact on the subsequent developmental processes, makes it central to our understanding of early embryonic development.
The difference between primary and secondary induction is a difference between
Neighboring cells influence the development of each other,either by direct contact or by production of chemical signals,in
The major group of the animal kingdom that includes animals that in their development show spiral cleavage and mosaic development is the __________.
Discuss or outline the derivatives of endoderm,mesoderm,and ectoderm.
Somites,which form segmental muscles and vertebrae,develop from which germ layer?
Eggs with abundant yolk that is concentrated at the vegetal pole are called
The concept of ________ suggested that preformed animals in the egg unfolded during development.
The conclusion of Spemann's work,in which he tied off a zygote isolating the nucleus on one side and only cytoplasm on the other,helped prove
A true coelom is a fluid-filled cavity completely lined by ___________.
After the fusion of sperm and egg membranes,there is the release of enzyme-rich granules from the egg that ultimately causes the separation of the vitelline envelope and the egg membrane;this is called
The special blastomere labeled the 4-d cell often gives rise to the
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